Many sagas tell of Leman Russ, Lord of Winter and War, the most formidable of the ghosts - the sons of the Emperor. At the peak of the Great Crusade, the Space Wolves attempt to bring the rebellious planet Doolan to Harmony. Enraged by the insubordination of the tyrant Darat, Russ vows to personally execute the ruler, but Lev El'Johnson, Lehmann's brother and master of the Dark Angels, advises to act more cautiously. Now the troops of Darat are opposed by two legions of Astartes, but tensions between the allies are growing, and the rivalry of the Wolf and the Lion threatens to overwhelm everyone.